north american species of cactus(北美仙人掌)-第2部分
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material。The whole group is to be regarded as made up of poorly
differentiated forms and only long observation under cultivation can
determine the possibilities of specific variation under the influence of
environment； of age； of inherent tendencies。For instance； that these plants
change in form and in spine characters with increasing age and after they
have begun to flower can not be doubted； but what described forms have
thus been separated in descriptions can only be guessed at。
John M。 Coulter。Lake Forest University；Lake Forest； Ill。； January；
1891。
CACTUS； ANHALONIUM； AND
LOPHOPHORA。
1。 CACTUS Linn。 Sp。 Pl。 466 （1753）； restricted。
MAMILLARIA Haw。 Synop。 177 （1812）； not Stackh。 （1809）。
Usually globose to oblong plants （simple； branching or cespitose）； but
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North American Species of Cactus
sometimes slender…cylindrical； covered with spine…bearing tubercles：
flower…bearing areola axillary （with reference to tubercles）； entirely
separate from the terminal spine…bearing areola； although sometimes
（Coryphantha） connected with it by a woolly groove along the upper face
of the tubercle： ovary naked： seeds smooth or pitted： embryo usually
straight； with short cotyledons。 Originally defined by Linnaeus in his
Systema； ed。 l （1735）。
The Linnaean genus Cactus of 1753 included 22 species and was
coextensive with the present order。In 1812 the species were separated by
Haworth into five genera； the original generic name Cactus being
discarded。Among these species C。 mamillaris seems to have stood as the
type； not only of the Linnaean genus Cactus； but also of Haworth's
Mamillaria； and as such should retain the original generic name。Besides；
Mamillaria was used as the generic name of an alga in 1809。Cactus
mamillaris L。 is the West Indian Mamillaria simplex Haw。
From one point of view the two sections of the genus （Eumamillaria
and Coryphantha） deserve generic separation； for the character of
grooveless and grooved tubercles seems to hold without exception； and the
sections are separated with more certainty than are certain species of
Coryphantha and Echinocactus。If genera are simply groups of
convenience the separation should be made。
I。 EUMAMILLARIA。Flowers from the axils of the older or full…grown
tubercles （hence usually appearing lateral）； mostly small； and generally
from whitish to pink or red： tubercles never grooved： fruit almost always
clavate and scarlet。
A。 Tubercles more or less quadrangular。
*Central spines not hooked。 ＋More than one central spine。
1。 Cactus alternatus； sp。 nov。
Subglobose； 10 cm。 in diameter； simple： tubercles long （15 to 20 mm。）
and spreading； with woolly axils： radial spines 3； rigid and recurved； 5 mm。
long； central spines 3； very stout and much recurved； 20 to 30 mm。 long；
alternating with the radials； all ashy colored and often twisted： flower and
fruit unknown。Type in Herb。 Coulter。
The few spines； with the very short radials alternating with the very
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North American Species of Cactus
long and stout centrals； furnish a striking character。 Occasionally one of
the centrals is wanting。
2。Cactus acanthophlegmus （Lehm。） Kuntze； Rev。 Gen。 Pl。 260
（1891）。
Mamillaria acanthophlegma Lehm。 Delect。 Sem。 Hamb。 （1833）
Subglobose with a deeply depressed vertex； or becoming cylindrical； 3
to 8。5 cm。 in diameter： tubercles sharply quadrangular…conical； with
densely woolly axils： radial spines 15 to 30； white； very slender （bristly）
and radiant； sometimes coarse capillary； 4 to 7 mm。 long； interwoven with
those of neighboring tubercles and so covering the whole plant； central
spines 2 to 4； robust and straight； erect or divergent； whitish or reddish；
black…tipped； 5 to 6。5 mm。 long： flowers reddish； 1 to 2 cm。 broad： fruit
unknown。Type unknown。
From Coahuila and San Luis Potosi to Oaxaca。Fl。 May。
Specimens examined： Coahuila （Poselger of 1856； Pringle 3116 of
1890）： San Luis Potosi （Eschanzier of 1891）。
The central spines are quite variable in number and arrangement。 In
case there are two they are vertically placed and are either erect and
parallel or widely divergent。Even three centrals may occur in the same
vertical plane； but more usually the three or four centrals are arranged
about a center and are widely divergent。The tubercles are apt to persist and
to become naked and corky with age。The axillary wool and the capillary
radials are also apt to be more or less persistent； thus giving the whole
plant a woolly appearance。
3。Cactus brandegei； sp。 nov。
Cylindrical： tubercles sharply quadrangular…conical； 6 to 8 mm。 long；
with densely woolly axils： radial spines about 10； slender and rigid；
whitish with dusky tips； spreading but not radiant； 7 to 10 mm。 long；
central spines 3 or 4； stouter and slightly longer； erect…spreading
（sometimes slightly curved）； reddish…brown below； becoming blackish
above： flowers small （scarcely longer than the tubercle？）： fruit
unknown。Type in Herb。 Calif。 Acad。
San Jorge； Lower California。Fl。 April。
Specimens examined： Lower California （Brandegee of 1889； at San
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Jorge）。
The species has somewhat the spine characters of C。 palmeri； but the
sharply quadrangular and longer tubercles with axillary wool free from
bristles suggest a very different affinity。
4。Cactus densispinus； sp。 nov。
Globose； 7。5 cm。 in diameter； simple： tubercles short； with woolly
axils： radial spines about 25； erect…spreading； slender but rigid； yellow
（brownish to black with age）； unequal； 8 to 10 mm。 long； central spines 6；
a little longer （10 to 12 mm。） and straight； more rigid and darker； black…
tipped： seeds obovate； reddish…brown； 1 mm。 long。Type in Herb。 Coulter。
Very easily distinguished by its dense； erect spines； which so
completely cover the plant as to give it the appearance of a large chestnut
bur。Another much smaller form； which seems to be a variety； has stouter
and longer ashy…white spines； the centrals darker…tipped； and the lower
centrals slightly curved。
＋＋ One short central spine （rarely two or none）： ovaries immersed：
seeds small； yellow and rugulose： simple。
5。Cactus heyderi （Muhlenpf。） Kuntze； Rev。 Gen。 Pl。 260 （1891）。
Mamillaria heyderiMuhlenpf。 Allg。 Gart。 Zeit。 xvi。 20（1848）。
Mamillaria declivisDietr。 Allg。 Gart。 Zeit。 xviii。 235 （1850）。 Mamillaria
applanataEngelm。 Pl。 Lindh 198 （1850）。Mamillaria texensisLabouret；
Monogr。 Cact。 89 （1858）。
Depressed； globose； usually with depressed vertex； 8 to 12 cm。 broad；
2。5 to 5 cm。 high： tubercles elongated： radial spines 10 to 22； whitish； 5 to
12 mm。 long； the lower usually the longer； stouter； and often darker；
central spine 4 to 8 mm。 long； light yellowish…brown； stout； straight； and
porrect： flowers 2 to 2。5 cm。 long； reddish…white： fruit incurved； 1。5 to 3
cm。 long。 （Ill。 Cact。 Mex。 Bound。 t。 9。 figs。 4…14）。Type unknown。
From the Guadalupe River； Texas； to the mouth of the Rio Grande； and
westward to Arizona and Sonora。Fl。 April； May。
Specimens examined： Texas （Lindheimer of 1845； 1847； 1853； Wright
226； also collections of 1849； 1852； 1853； 1855； 1856； Bigelow of 1853；
Trelease of 1892； Nealley of 1892）： New Mexico （Wright 311； Bigelow of
1853； Evans of 1891）： Arizona （Pringle of 1881）： also growing in Mo。 Bot。
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North American Species of Cactus
Gard。 1893； and in the World's Fair collection of Mrs。 Nickels。
The radial spines are somewhat variable in relative length； often
becoming almost equal； while sometimes the upper radials are very much
reduced。The figure referred to in Cact。 Mex。 Bound。 is not satisfactory as
to the general habit of the plant； which is flat…topped rather than
hemispherical。
6。Cactus heyderi hemisphaericus （Engelm。）。
Mamillaria hemisphaericaEngelm。 Pl。 Lindh。 198 （1850）。
Differs in being hemispherical instead of flat…topped； in its fewer （9 to
12） and shorter （4 to 8 mm。） radial spines； and much smaller less rough
and lighter…colored seeds。（Ill。 Cact。 Mex。 Bound。 t。 9。 figs。 15…17）Type； the
〃Goebel's Garden〃 plants in Herb。 Mo。 Bot。 Gard。
Throughout southern Texas and southern New Mexico； and southward；
not extending so
