autobiography and selected essays-第23部分
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who has watched its display; continued hour after hour; without
pause or sign of weakening。 The possible complexity of many other
organic forms; seemingly as simple as the protoplasm of the nettle;
dawns upon one; and the comparison of such a protoplasm to a body
with an internal circulation; which has been put forward by an
eminent physiologist; loses much of its startling character。
Currents similar to those of the hairs of the nettle have been
observed in a great multitude of very different plants; and weighty
authorities have suggested that they probably occur; in more or
less perfection; in all young vegetable cells。 If such be the
case; the wonderful noonday silence of a tropical forest is; after
all; due only to the dulness of our hearing; and could our ears
catch the murmur of these tiny Maelstroms; '96' as they whirl in the
innumerable myriads of living cells which constitute each tree; we
should be stunned; as with the roar of a great city。
Among the lower plants; it is the rule rather than the exception;
that contractility should be still more openly manifested at some
periods of their existence。 The protoplasm of Algae and Fungi
becomes; under many circumstances; partially; or completely; freed
from its woody case; and exhibits movements of its whole mass; or
is propelled by the contractility of one; or more; hair…like
prolongations of its body; which are called vibratile cilia。 And;
so far as the conditions of the manifestation of the phaenomena of
contractility have yet been studied; they are the same for the
plant as for the animal。 Heat and electric shocks influence both;
and in the same way; though it may be in different degrees。 It is
by no means my intention to suggest that there is no difference in
faculty between the lowest plant and the highest; or between plants
and animals。 But the difference between the powers of the lowest
plant; or animal; and those of the highest; is one of degree; not
of kind; and depends; as Milne…Edwards '97' long ago so well pointed
out; upon the extent to which the principle of the division of
labour is carried out in the living economy。 In the lowest
organism all parts are competent to perform all functions; and one
and the same portion of protoplasm may successfully take on the
function of feeding; moving; or reproducing apparatus。 In the
highest; on the contrary; a great number of parts combine to
perform each function; each part doing its allotted share of the
work with great accuracy and efficiency; but being useless for any
other purpose。
On the other hand; notwithstanding all the fundamental resemblances
which exist between the powers of the protoplasm in plants and in
animals; they present a striking difference (to which I shall
advert more at length presently); in the fact that plants can
manufacture fresh protoplasm out of mineral compounds; whereas
animals are obliged to procure it ready made; and hence; in the
long run; depend upon plants。 Upon what condition this difference
in the powers of the two great divisions of the world of life
depends; nothing is at present known。
With such qualifications as arises '98' out of the last…mentioned
fact; it may be truly said that the acts of all living things are
fundamentally one。 Is any such unity predicable of their forms?
Let us seek in easily verified facts for a reply to this question。
If a drop of blood be drawn by pricking one's finger; and viewed
with proper precautions; and under a sufficiently high microscopic
power; there will be seen; among the innumerable multitude of
little; circular; discoidal bodies; or corpuscles; which float in
it and give it its colour; a comparatively small number of
colourless corpuscles; of somewhat larger size and very irregular
shape。 If the drop of blood be kept at the temperature of the
body; these colourless corpuscles will be seen to exhibit a
marvellous activity; changing their forms with great rapidity;
drawing in and thrusting out prolongations of their substance; and
creeping about as if they were independent organisms。
The substance which is thus active is a mass of protoplasm; and its
activity differs in detail; rather than in principle; from that of
the protoplasm of the nettle。 Under sundry circumstances the
corpuscle dies and becomes distended into a round mass; in the
midst of which is seen a smaller spherical body; which existed; but
was more or less hidden; in the living corpuscle; and is called its
nucleus。 Corpuscles of essentially similar structure are to be
found in the skin; in the lining of the mouth; and scattered
through the whole framework of the body。 Nay; more; in the
earliest condition of the human organism; in that state in which it
has but just become distinguishable from the egg in which it
arises; it is nothing but an aggregation of such corpuscles; and
every organ of the body was; once; no more than such an
aggregation。
Thus a nucleated mass of protoplasm turns out to be what may be
termed the structural unit of the human body。 As a matter of fact;
the body; in its earliest state; is a mere multiple of such units;
and in its perfect condition; it is a multiple of such units;
variously modified。
But does the formula which expresses the essential structural
character of the highest animal cover all the rest; as the
statement of its powers and faculties covered that of all others?
Very nearly。 Beast and fowl; reptile and fish; mollusk; worm; and
polype; are all composed of structural units of the same character;
namely; masses of protoplasm with a nucleus。 There are sundry very
low animals; each of which; structurally; is a mere colourless
blood…corpuscle; leading an independent life。 But; at the very
bottom of the animal scale; even this simplicity becomes
simplified; and all the phaenomena of life are manifested by a
particle of protoplasm without a nucleus。 Nor are such organisms
insignificant by reason of their want of complexity。 It is a fair
question whether the protoplasm of those simplest forms of life;
which people an immense extent of the bottom of the sea; would not
outweigh that of all the higher living beings which inhabit the
land put together。 And in ancient times; no less than at the
present day; such living beings as these have been the greatest of
rock builders。
What has been said of the animal world is no less true of plants。
Imbedded in the protoplasm at the broad; or attached; end of the
nettle hair; there lies a spheroidal nucleus。 Careful examination
further proves that the whole substance of the nettle is made up of
a repetition of such masses of nucleated protoplasm; each contained
in a wooden case; which is modified in form; sometimes into a woody
fibre; sometimes into a duct or spiral vessel; sometimes into a
pollen grain; or an ovule。 Traced back to its earliest state; the
nettle arises as the man does; in a particle of nucleated
protoplasm。 And in the lowest plants; as in the lowest animals; a
single mass of such protoplasm may constitute the whole plant; or
the protoplasm may exist without a nucleus。
Under these circumstances it may well be asked; how is one mass of
non…nucleated protoplasm to be distinguished from another? why call
one 〃plant〃 and the other 〃animal〃?
The only reply is that; so far as form is concerned; plants and
animals are not separable; and that; in many cases; it is a mere
matter of convention whether we call a given organism an animal or
a plant。 There is a living body called Aethalium septicum; which
appears upon decaying vegetable substances; and; in one of its
forms; is common upon the surfaces of tan…pits。 In this condition
it is; to all intents and purposes; a fungus; and formerly was
always regarded as such; but the remarkable investigations of De
Bary '99' have shown that; in another condition; the Aethalium is an
actively locomotive creature; and takes in solid matters; upon
which; apparently; it feeds; thus exhibiting the most characteristic
feature of animality。 Is this a plant; or is it an animal?
Is it both; or is it neither? Some decide in favour of the last
supposition; and establish an intermediate kingdom; a sort
of biological No Man's Land '100' for all these questionable forms。
But; as it is admittedly impossible to draw any distinct boundary line
between this no man's land and the vegetable world on the one hand;
or the animal; on the other; it appears to me that this proceeding
merely doubles the difficulty which; before; was single。
Protoplasm; simple or nucleated; is the formal basis of all life。
It is the clay of the potter: which; bake it and paint it as he
will; remains clay; separated by artifice; and not by nature; from
the commonest brick or sun…dried clod。
Thus it becomes clear that all living powers are cognate; and
that all living forms are fundamentally of one character。 The
researches of the chemist have revealed a no less striking
uniformity of material composition in living matter。
In perfect strictness; it is true that chemical investigation can
tell us little or nothing; directly; of the composition of living
matter; inasmuch as such matter must needs die in the act of
analysis;and upon this very obvious ground; objections; which I
confess seem to me to be somewhat frivolous; have been raised to
the drawing of any conclusions whatever respecting the composition
of actually living matter; from that of the dead matter of life;
which alone is accessible to us。 But objectors of this class do
not seem to reflect that it is also; in strictness; true that we
know nothing about the composition of any body whatever; as it is。
The statement that a crystal of calc…spar consists of carbonate of
lime; is quite true; if we only mean that; by appropriate
processes; it may be resolved into carbonic acid and quicklime。 If
you pass the same carbonic acid over th